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An 8-year-old boy is seen by an ophthalmologist for vision difficulties, and the physician notices a slowing of the boy’s eye movements. The ophthalmologist finds ophthalmoplegia and pigmentary retinopathy and suspects the child has Kearns-Sayre syndrome. Assuming that the defect in this disorder is due to a mutation in complex II of the ETC, electron transfer from which substrate would be impaired?